Genus:

Eremias FITZINGER, 1834

Species (45):

Eremias acutirostris  (BOULENGER, 1887)

Eremias afghanistanica  BÖHME & SZCZERBAK, 1991

Eremias andersoni  DAREVSKY & SZCZERBAK, 1978

Eremias argus  PETERS,1869

Eremias arguta  (PALLAS, 1773)

Eremias aria  ANDERSON & LEVITON, 1967

Eremias brenchleyi  GÜNTHER, 1872

Eremias buechneri  BEDRIAGA, 1907

Eremias cholistanica  BAIG & MASROOR, 2006

Eremias dzungarica  ORLOVA et al., 2017

Eremias fahimii  MOZAFFARI & AHMADZADEH & SABERI-PIROOZ, 2020

Eremias fasciata  BLANFORD, 1874

Eremias grammica  (LICHTENSTEIN, 1823)

Eremias graphica  ORLOVA, E. RASEGAR-POUYANI, RAJABIZADEH, NABIZADEH, POYARKOV, MELNIKOV & NAZAROV, 2023

Eremias intermedia  (STRAUCH, 1876)

Eremias isfahanica  RASTEGAR-POUYANI et al., 2016

Eremias kakari  MASROOR et al., 2020

Eremias kavirensis  MOZAFFARI & PARHAM, 2007

Eremias killasaifullahi  MASROOR et al., 2022

Eremias kokshaaliensis  EREMCHENKO & PANFILOV 1999

Eremias kopetdaghica  SZCZERBAK, 1972

Eremias lalezharica  MORAVEC, 1994

Eremias lineolata  (NIKOLSKY, 1896)

Eremias montana  RASTEGAR-POUYANI & RASTEGAR-POUYANI, 2001

Eremias multiocellata  GÜNTHER, 1872

Eremias nigrocellata  NIKOLSKY, 1896

Eremias nikolskii  BEDRIAGA in NIKOLSKY, 1905

Eremias papenfussi  MOZAFFARI, AHMADZADEH & PARHAM, 2011

Eremias persica  BLANFORD, 1874

Eremias pleskei  BEDRIAGA, 1905

Eremias przewalskii  (STRAUCH, 1876)

Eremias pseudofasciata  ORLOVA, E. RASEGAR-POUYANI, RAJABIZADEH, NABIZADEH, POYARKOV, MELNIKOV & NAZAROV, 2023

Eremias quadrifrons  (STRAUCH, 1876)

Eremias rafiqi  MASROOR et al., 2022

Eremias regeli  BEDRIAGA in NIKOLSKY, 1905

Eremias roborowskii  (BEDRIAGA, 1912)

Eremias scripta  (STRAUCH, 1867)

Eremias sp. 

Eremias strauchi  KESSLER, 1878

Eremias stummeri  WETTSTEIN, 1940

Eremias suphani  BAŞOĞLU & HELLMICH, 1968

Eremias szczerbaki  JERIOMTSCHENKO & PANFILOV, 1992

Eremias velox  (PALLAS, 1771)

Eremias vermiculata  BLANFORD, 1875

Eremias yarkandensis  BLANFORD, 1875

Species Typica:

Eremias velox  (PALLAS, 1771)

Taxonomic notes:

Die ehemalige Sammelgattung Eremias wurde von SHCHERBAK (1971) in verschiedene Gattungen aufgeteilt, indem er die `sections` BOULENGER`s (1921) in Gattungsrang erhob, aber die paläarktischen Arten von Scapteira mit Eremias vereinigte. Die nun rein paläarktische Gattung teilte er in fünf Untergattungen (Eremias, Ommateremias, Pareremias, Rhabderemias, Scapteira). Nach EREMCHENKO (1999) ist die species typica der Gattung nicht Lacerta velox (= Eremias velox) sondern Lacerta variabilis (= Eremias arguta), weshalb er nun die ehemalige Ommateremias als Eremias führt und die velox-Gruppe in eine neue Untergattung Dimorphea stellt. BARABANOV & ANDREI fanden für die selbe Gruppe den Namen Aspidorhinus EICHWALD 1841 als verfügbar. Da die entsprechenden Publikationen aber in gepflegtem Russisch abgefasst und die Argumentationen somit schwer nachvollziehbar sind, werden im folgenden die Untergattungen weiter mit den von SHCHERBAK eingeführten Namen geführt.
Ähnlich wie bei Acanthodactylus wurden in den letzten Jahren zahlreiche Arten beschrieben, deren Validität in vielen Fällen zu Zweifeln Anlass gibt. Trotzdem werden hier vorderhand alle nominellen Arten (kritiklos) aufgeführt. Eine umfassende genetische Analyse der Gattung wäre in höchstem Ausmaß wünschenswert, ist bislang aber ausständig.

WERNER MAYER, 2015

Relevant taxonomic literature:

• Wiegmann, A.F.A. (1834) -  Familia 1: Lacertidae -  Herpetologia mexicana, seu Descriptio amphibiorum Novae Hispaniae quae itineribus comitis De Sack, Ferdinandi Deppe et Chr. Guil. Schiede in Museum zoologicum Berolinense pervenerunt. Pars prima saurorum species amplectens, adjecto systematis saurorum prodromo, additisque multis in hunc amphibiorum ordinem observationibus. C. G. Luderitz, Berolini: 9-10.  

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• Boulenger, G.A. (1921) -  Monograph of the Lacertidae. Vol. II. -  British Museum (Natural History). Department of Zoology. London. 451 pp.  

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• Szczerbak, N.N. (1971) -  Ecology of the Gobi Racerunner (Eremias przewalski str.). -  Vestnik zoologii, Journal of the Schmalhausen Institute of Zoology, National Academy of Sciences of Ukraine, Kiev, 1971 (4): 58-66.  

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• Eremchenko, V.K. & Panfilov A.M. (1999) -  Taxonomic position and geographic relations of a lacertid lizard Eremias velox from the Issyk-Kul lake depression, Tien Shan mountains, Kyrgyzstan.  -  Nauka i Novye Tekhnologii, Seriya Biologicheskaya, 1: 119-125.  

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• Barabanov, A.V. (2009) -  Aspidorhinus Eichwald, 1841 as a valid subgeneric name for Eremias velox species group (Sauria, Lacertidae). -  Current Studies in Herpetology, Saratov, Russia, 9 (1-2): 59-61.  

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  Some nomenclature problems of the Eremias (Sauria, Lacertidae) genus are discussed. Aspidorhinus Eichwald, 1841 is shown to be a senior synonym of Dimorphea Eremchenko, 1999 and so should be a valid subgeneric name for the Eremias velox species group. A list of the subgenus’ species is given.

  Барабанов А.В. (2009) -  Aspidorhinus Eichwald, 1841 - валидное подродовое название для видовой группы Eremias velox (Sauria, Lacertidae). -  Современная герпетология, 9 (1-2): 59-61.    

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  Обсуждаются некоторые номенклатурные проблемы рода Eremias. Показано, что Dimorphea Eremchenko, 1999 - младший синоним Aspidorhinus Eichwald, 1841, поэтому именно последнее название следует использовать в качестве валидного подродового для группы видов Eremias velox. Приведен список видов, входящих в данный подрод.

• Guo, X. & Dai, X. & Chen, D. & Papenfuss, T.J. & Ananjeva, N.B. & Melnikov, D.A. & Wang, Y. (2011) -  Phylogeny and divergence times of some racerunner lizards (Lacertidae: Eremias) inferred from mitochondrial 16S rRNA gene segments. -  Molecular Phylogenetics and Evolution, 61 (2): 400–412.    

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  Eremias, or racerunners, is a widespread lacertid genus occurring in China, Mongolia, Korea, Central Asia, Southwest Asia and Southeast Europe. It has been through a series of taxonomic revisions, but the phylogenetic relationships among the species and subgenera remain unclear. In this study, a frequently studied region of the mitochondrial 16S rRNA was used to (i) reassess the phylogenetic relationships of some Eremias species, (ii) test if the viviparous species form a monophyletic group, and (iii) estimate divergence time among lineages using a Bayesian relaxed molecular-clock approach. The resulting phylogeny supports monophyly of Eremias sensu Szczerbak and a clade comprising Eremias, Acanthodactylus and Latastia. An earlier finding demonstrating monophyly of the subgenus Pareremias is corroborated, with Eremias argus being the sister taxon to Eremias brenchleyi. We present the first evidence that viviparous species form a monophyletic group. In addition, Eremias przewalskii is nested within Eremias multiocellata, suggesting that the latter is likely a paraphyletic species or a species complex. Eremias acutirostris and Eremias persica form a clade that is closely related to the subgenus Pareremias. However, the subgenera Aspidorhinus, Scapteira, and Rhabderemias seem not to be monophyletic, respectively. The Bayesian divergence-time estimation suggests that Eremias originated at about 9.9 million years ago (with the 95% confidence interval ranging from 7.6 to 12 Ma), and diversified from Late Miocene to Pleistocene. Specifically, the divergence time of the subgenus Pareremias was dated to about 6.3 million years ago (with the 95% confidence interval ranging from 5.3 to 8.5 Ma), which suggests that the diversification of this subgenus might be correlated with the evolution of an East Asian monsoon climate triggered by the rapid uplift of the Tibetan Plateau approximately 8 Ma.